Cultivated tomato cannot produce pu

Cultivated tomato cannot produce purple fruits: flavonoid biosynthesis is interrupted with the accumulation of intermediate compounds (mainly naringenin chalcone and the flavonol glycosides rutin and kaempferol-3-O-rutinoside) (Bovy et al., 2002), probably due to an inefficient activation of the pathway (Povero et al., 2011).However, due to their increasingly recognized health-promoting effects (Martin et al., 2011; Liu et al., 2017),considerable work has been done in recent years to enrich tomatoes with anthocyanins (Gonzali et a., 2009;Martì et al., 2016). Along with transgenic approaches (Butelli et al., 2008), biodiversity has been exploitedwith positive results (Mes et al., 2008; Gonzali et al., 2009).